Interesting links M, I didn't comment earlier as it all felt a bit baffling. You've probably seen this link which is related as about DNA testing again and also references your first link. This time about the origins of horses ... which is intriguing as horses appear on rock/cave paintings.
http://www.independent.co.uk/news/science/horse-origin-mysterious-domestic-dna-analysis-przewalskis-botai-domesticated-a8224121.html
See also the Peche Merle spotted horses - which seem to be dated between 25,000BC and 16,000BC
https://en.wikipedia.org/wiki/Pech_Merle
Read in conjunction with the Independent's article it all seems a bit confusing.
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